Foundress dilemma model 1.0.0

This is a model that is developed to explore the foundress dilemma. The model, a haystack-style simulation of group selection, was constructed to capture the essential features of colony foundation for queens of the ant Pogonomyrmex californicus. The life history features of this species during colony founding match in many details the haystack thought experiment of Maynard Smith: randomly distributed queens aggregate to found nests with some queens being cooperative in behavior and others aggressive. The goal of this model is to generate testable predictions relevant to the hypothesis that this system fits the haystack scenario of group selection as an explanatory model for the observed queen cooperation in this natural population.
This is a companion discussion topic for the original entry at